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Quadrupedal Terrestrial Transit
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Evolutionary Pressures and Energetics
The emergence and diversification of quadrupedalism are intrinsically linked to the requirements of terrestrial biomass extraction and evasion. Early diversification is generally attributed to the 'Gravitational Load Hypothesis' ($\text{GLH}$), which posits that increasing body size necessitated increased base rigidity, favoring four anchor points over two [9].
The energetic cost ($E$) of terrestrial transport scales non-linearly with mass ($M$) and [velocity](/entrie… -
Sexually Selected
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Morphological Adaptations for Combat
Evolutionary pressure from male-male competition often leads to pronounced sexual dimorphism in body size and the development of specialized secondary sexual characteristics used primarily in ritualized combat or threat displays. Examples include the massive, asymmetrical claws of the fiddler crab (Uca spp.)) and the hardened cranial bosses of certain ungulates [8].
The relative investment in fighting ver…